More samples from Golestan (around Gorgan on map below) show Indo-European (or maybe a different EHG-rich group?) presence by 3100 BC. Two had Y haplogroup J, one T1a. Authors believe they came by Central Asia rather than Caucasus. cell.com
Proto-Greek speakers are confirmed to have arrived in Greece during the 23rd century BC droughts in the eastern Mediterranean which also likely brought down Old Kingdom Egypt as well as a number of other states.
Two Greek finds are interesting - G23 from Theopetra (Thessaly) with Y haplogroup I & G31 from Perachora (Peloponnesus) with Y haplogroup BT. G23 has quite a bit of steppe ancestry, while G31 has none. BT is a sub-Saharan African haplogroup, while haplogroup I has WHG origin.
The BT might just be result of low genome coverage - it was only
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